Pterostichus (Nialoe) shotaroi kiimontanus
Phylogenetic studies of the subgenus Nialoe (s. lat.) (Coleoptera, Carabidae, genus Pterostichus), Part 2: the asymmetricus species group. Zoological Science (Tokyo), 22(11), November 2005: 1217-1228. 1220 [Zoological Record Volume 142]